Eaten by

• White-sided Dolphins (Lagenorhynchus)
• Typical Cormorants And Shags (Phalacrocorax)
• Neotropic Cormorant (Phalacrocorax brasilianus)
• Large White-headed Gulls (Larus)
• Royal Tern (Thalasseus maximus)
• Western Gull (Larus occidentalis)
• Brandt's Cormorant (Phalacrocorax penicillatus)
• Pelicans (Pelecanus)
• Red-footed Booby (Sula sula)
• Least Tern (Sternula antillarum)
• Elegant Tern (Thalasseus elegans)
• Brown Booby (Sula leucogaster)
• Common Murre (Uria aalge)
• Little Penguin (Eudyptula minor)
• Forster's Tern (Sterna forsteri)
• Heermann's Gull (Larus heermanni)
• Caspian Tern (Hydroprogne caspia)
• Small Shearwaters (Puffinus)
• Rhinoceros Auklet (Cerorhinca monocerata)
• Tufted Puffin (Fratercula cirrhata)
• Double-crested Cormorant (Phalacrocorax auritus)
• Peruvian Booby (Sula variegata)
• (Otaria flavescens)
• Spectacled Porpoise (Phocoena dioptrica)
• (Pelecanus thagus)
• Blue-footed Booby (Sula nebouxii)
• Humboldt Penguin (Spheniscus humboldti)
• (Phalacrocorax gaimardi)
• Inca Tern (Larosterna inca)
• (Puffinus griseus)
• Indian Cormorant (Phalacrocorax fuscicollis)
• (Pelecanus conspicillatus)
• (Puffinus tenuirostris)
• Masked Booby (Sula dactylatra)
• Heinroth's Shearwater (Puffinus heinrothi)
• Cape Cormorant (Phalacrocorax capensis)
• (Larus saundersi)
• Nazca Booby (Sula granti)
• African Penguin (Spheniscus demersus)
• (Phalacrocorax niger)
• Matsudaira's Storm-petrel (Oceanodroma matsudairae)
• Townsend's Shearwater (Puffinus auricularis)
• Band-rumped Storm-petrel (Oceanodroma castro)
• Peruvian Diving-petrel (Pelecanoides garnotii)
• Belcher's Gull (Larus belcheri)
• (Larus fuliginosus)
• (Oceanodroma markhami)
• (Phalacrocorax coronatus)
• (Phalacrocorax sulcirostris)
• (Pelecanus crispus)
• (Larus atricilla)
• Fairy Tern (Sternula nereis)
• Audubon's Shearwater (Puffinus lherminieri)
• (Thalasseus eurygnatha)
• (Larus novaehollandiae)
• Polynesian Storm-petrel (Nesofregetta fuliginosa)
• (Phalacrocorax albiventer)
• Wedge-rumped Storm-petrel (Oceanodroma tethys)
• Lesser Crested Tern (Thalasseus bengalensis)
• (Thalassarche carteri)
• Spot-billed Pelican (Pelecanus philippensis)
• Bounty Islands Shag (Phalacrocorax ranfurlyi)
• Yellow-billed Tern (Sternula superciliaris)
• White-fronted Tern (Sterna striata)
• Little Tern (Sternula albifrons)
• Stewart Island Shag (Phalacrocorax chalconotus)
• (Larus scopulinus)
• River Tern (Sterna aurantia)
• Saunders's Tern (Sternula saundersi)
• Japanese Cormorant (Phalacrocorax capillatus)
• Chatham Islands Shag (Phalacrocorax onslowi)
• Black-faced Cormorant (Phalacrocorax fuscescens)
• Flightless Cormorant (Phalacrocorax harrisi)
• Black Storm-petrel (Oceanodroma melania)
• Damara Tern (Sternula balaenarum)
• Cape Verde Shearwater (Calonectris edwardsii)
• Ashy Storm-petrel (Oceanodroma homochroa)
• (Sternula lorata)
• Pitt Island Shag (Phalacrocorax featherstoni)
• Great Cormorant (Phalacrocorax carbo)
• Spotted Shag (Phalacrocorax punctatus)
• Black-bellied Storm-petrel (Fregetta tropica)
• Greater Crested Tern (Thalasseus bergii)
• Swinhoe's Storm-petrel (Oceanodroma monorhis)
• (Puffinus pacificus)
• Brown Noddy (Anous stolidus)
• Noddies (Anous)
• (Puffinus carneipes)
• Magellanic Penguin (Spheniscus magellanicus)
• (Papasula abbotti)
• Yellow-footed Gull (Larus livens)
• (Ichthyaetus ichthyaetus)
• (Larus brunnicephalus)
• Australasian Gannet (Morus serrator)
• Galápagos Penguin (Spheniscus mendiculus)
• Mew Gull (Larus canus)
• Black-fronted Tern (Chlidonias albostriatus)
• Snowy-crowned Tern (Sterna trudeaui)
• Cape Gannet (Morus capensis)
• White-cheeked Tern (Sterna repressa)

Preyed upon by

• Double-crested Cormorant (Phalacrocorax auritus)
• Inca Tern (Larosterna inca)